Runejl,
Pre-dam, there was a natural barrier to salmonid migration a short distance upstream of present day Diablo Dam. Major cataracts exist between there and Ruby Creek, which is just upstream of Ross Dam. Salmon and steelhead migrated upstream of present day Gorge Dam and spawned on Cedar Bar and Reflector Bar and in the lower two or three miles of Stetattle Creek. During the Skagit relicensing proceeding years ago, I calculated the amount of salmon (chinook) and steelhead habitat lost. I don't remember the value, but it was small, all things considered, and is mitigated for by having Seattle provide improved spawning and incubation flows downstream of Gorge Dam.
The presence of rainbow trout and native char upstream of Ross, and the historic records documenting them, indicate that passage through the cataract has been possible, even if temporarily, at times during the historic past. A former WDFW biologist asserted that summer steelhead formerly populated the upper Skagit. They may have, but not in the historic past, or time frame that would affect Seattle City Light's mitigation obligations. The upper Skagit Valley upstream of the Ross Dam site was settled prior to dam construction. There were a couple of horse ranchers upstream of Ruby Creek, along with a few trappers and miners. They caught trout and char, but no salmon or steelhead. Skagit Indians who traveled to a mining site where they made arrowheads and tools packed their own dried salmon with them from downstream, knowing they would not be able to catch any fresh salmon up there.
It wouldn't be fair to expect anyone from Sedro Woolley to ever believe folks from Seattle, so it's the good thing that the above information comes from more reliable sources. Those sources are from Indian oral tradition stories and former residents of the upper valley who were pissed at having to move out because Seattle's dam flooded their land. People who were not from Seattle or benefitting from Seattle seem like pretty good sources.
The Baker River dams flooded most of the productive salmon and steelhead reaches of the Baker sub-basin, extirpating native chinook and steelhead, and nearly wiping out the coho and sockeye until restoration measures began in 1985. The six miles of remaining accessible salmon habitat upstream of Baker Lake is damn close to being sterile water, and therefore not too productive. Bull trout and some salmon do spawn there however. Because of the recent relicensing of the Baker dams, Puge Sound Energy (PSE) is making significant improvements to upstream and downstream fish passage around the dams. PSE is also building a new "small scale" hatchery at the upper end of Lake Shannon. It's primary focus is on enhancement of the Baker sockeye population, but it includes facilities so the managers can attempt restoration of chinook and steelhead also, should they decide to. Coho are doing pretty well with the new fish passage system, and the small amount of hatchery supplementation of coho may be discontinued if the natural population is self-sustaining.
Sg